Ronald Altig1, Roy W. McDiarmid2, Kimberly A. Nichols3 and Paul C. Ustach4
SECTION 1. ORAL APPARATUS AND RANGE: Division 1, Sections 1-A and 2-A - [5] -
A. | medial margins of oral flaps divergent and with smooth edges; temporary pools during summer; widespread east and west of Mississippi River |
Gastrophryne - Section 2
B. | medial margins of oral flaps convergent and papillate or crenulate; temporary pools with seasonal rains; extreme southern Texas (sympatric: Gastrophryne olivacea and Rhinophrynus dorsalis) |
Hypopachus variolosus [B]
SECTION 2. COLORATION AND RANGE: Division 1, Sections 1-A and 2-A; Division 5, Section 1-A - medial margins of oral flaps divergent and with smooth edges; west of Mississippi River - [3] -
A. | dorsum mostly black in life (lighter at night) and preservative, never gray or brassy; belly dark with pale blotches and mottling; lateral pale tail stripe bright and distinct at least basally; temporary lentic sites formed by seasonal rains; eastern Texas north to central Missouri (sympatric: Gastrophryne olivacea*) |
Gastrophryne carolinensis (part) [A]
B. | dorsum olive-gray to brown to brassy caused by profuse iridophore pigmentation in life; venter white or coppery in life; lateral tail stripe faint to absent; temporary lentic sites formed by seasonal rains; most of Nebraska south to southern Texas plus south-central Arizona (sympatric: Gastrophryne carolinensis* and Rhinophrynus dorsalis) |
Gastrophryne olivacea [B]